| In recent years, along with exacerbation of global eco-environmental problem, ecological function of vegetation restoration and reconstruction has been one of ecological hotspots regarded by people. Moreover, light stress, drought and rise of atmosphere CO2 concentration are important ecological factors, which influence distributing and growth of vegetation. Photosynthetic gas exchange parameters has been researched on crop and economic forest centralized mostly. Research on variety rule of photosynthesis physiological index of Lianas was rather little under a series of soil moisture grads. Therefore, using , synthetically the theory of water transport of soil-plant-atmosphere continuum interface and theory of plant water use efficiency, and regarding Parthenocissus quinquefolia,Parthenocissus semicordata,Euonymus fortunei,Euonymus fortunei minimus Rhed,Campsis radicans Seen,Ampelopsis brevipedunculata,Wisteria sinensis Sweet,Lonicera japonica Thunb,Euonymus scandens Graham,Hedera neaplensis var. Sinensis et al. as experimental materials, water transport process and water physiological saving mechanism of leaf-atmosphere interface of woodiness Lianas has been studied. Thereby, quantitative relation among water transport efficiency of leaf-atmosphere interface, soil water content of root-soil interface and quantum flux of leaf-atmosphere interface are established, and stomata limit mechanism of soil moisture in root-soil interface driving WUE of leaf-atmosphere interface and chlorophyll fluorescence kinetic mechanism are explored. These theoretical and technical gists are very useful at selecting and deploying reasonable sites for eximious Lianas, in the process of vegetation restoration and reconstruction of droughty and barren hills. The results as follow:(1) The response of leaves photosynthetic efficiency to the dynamic effecters of soil moistureThe net photosynthetic rate (Pn),transpiration rate (Tr) and water use efficiency (WUE) of Lianas leaves have notable threshold response value to the level of soil moisture and the variation of photosynthetic active radiation (PAR). We can ascertain some parameters for maintaining the natural growth and water efficient use of Lianas, such as : the maximal soil moisture deficit, the upper limit and fitting soil water content, the upper,lower limit and fitting value of PAR. For example, W. sinensis:①In order to maintain the high-level Pn and WUE of W. sinensis ,synchronously, the fitting mass water content (Wm) is about 13.1%~22.6%, relative water content (Wr) is about 46.5%～80.6%, and the optimum Wm is about19.9% (Wr is 70.9%).②The adaptability of W. sinensis to light conditions is very fargoing, and the high-level Pn and WUE is appeared in the PAR of 600～1600μmol·m-2·s-1, and the light saturation points of Pn and WUE are all about 800～1000μmol·m-2·s-1.③The most deficient degree to Wm is 10.2% (Wr is 36.2%) for the normal growth of W. sinensis, in this situation, the leaf photosynthesis organ will be badly destroyed if the number of PAR exceeds 1000μmol·m-2·s-1, and it is the critical point of destructivity in leaf photosynthesis organ.④Establishing the response model of water transport efficiency of the interface between leaf-atmosphere to PAR, we find that the non-rectangular hyperbola model is very good to simulate light-response curve, and the convexity is about 1.With the increase of soil mass water content (Wm is about 5.5%~19.4%),relative water content(Wr) is about20.1%~71.1%, the number of light compensation point is decline while light saturation point,the maximum Pn,apparent quantity yield are all increase. When Wm is about 19.4%, the light compensation point is at the minimum of 21.61μmol·m-2·s-1, and light saturation point is at the maximum of 1400μmol·m-2·s-1.⑤With the aggravating of soil moisture stress, the reason of Pn'decline is changed from stomatal limit to No-stomatal limit. For example, the Lianas of W. sinensis, the appearance of No-stomatal limit is nearly correlative with soil moisture and light intensity. The decline of Pn is caused mainly by stomatal limit in the range of Wm between 15.7% and 22.6%, in this situation; the Pn is not affected obviously by PAR. Out of this soil moisture situation, the Pn is affected obviously by PAR, and the critical turning point of PAR is appeared with the change from stomatal limit to No- stomatal limit.(2) The response of daily trends of leaves photosynthetic efficiency under soil moisture stress ①The primary factors influencing Pn of H. neaplensis var. can be sum up two species, one is air temperature and relative humidity, the other is light intensity under abundant soil moisture. One is light intensity and atmosphere CO2 concentration, the other is air relative humidity under severe soil moisture stress.②On condition of all kinds of soil moisture, Pn of Lonicera japonica Thunb., especially the kind of L. j. cv.ungulata and L.j.cv. Variegatum,are correspondingly higher. On condition of moderate soil moisture stress, Pn of E. scandens,A. brevipedunculata,E. fortunei are correspondingly higher. Pn of Campsis radicans,E.f.var.minimus Rhed,P. quinquefolia,P. semicordata,Wisteria sinensis et al. are secondly higher. Compared with else Lianas, Pn of W. sinensis,P. quinquefolia,H. neaplensis var are lower although soil moisture condition is good.③With aggravation of soil moisture stress, transpiration rate and water use efficiency of most Lianas represents descendant trend and the variety of factors influencing transpiration rate is rather complex. From compositive factors in single factor's relativity analysis and stepwise regression equation, the influence of environmental factors on water use efficiency of C. radicans is rather complex under soil moisture stresss. Commonly, 4~5 environmental factors influence together water use efficiency of C. radicans, especially atmosphere CO2 concentration,intercellular CO2 concentration,leaf temperature.④With aggravation of soil moisture stress, the order of Lianas'cluster frequency of Pn and Tr is high mean>middling mean>low mean. The high,middling mean amount of breed is larger under check and mild soil moisture stress, and the low mean amount of breed is larger under moderate and severe soil moisture stress.(3) The response of leaves photosynthetic efficiency on CO2 concentration under different soil moisture①The increase of soil moisture is favorable for Lianas'photosynthesis under high CO2 concentration, and accelerates the use of high CO2 concentration.②With the aggravation of soil moisture stress,the CO2 saturation point of Lianas is reduced, and the change of the response curve is very mild under high CO2 concentration.With the aggravation of soil moisture stress,the frequency of low Pn value is increase for all Lianas under every CO2 concentration. ③With the aggravation of soil moisture stress, some Lianas'carboxylation efficiency is reduced, and the CO2 compensation points of H. neaplensis var,P. semicordata,W. sinensis,L. j. cv.tomentosa,L. j.cv. Variegatum et al, Lianas is increased, and theГmean of other Lianas is almost equal under mild and moderate soil moisture stress. The maximal regeneration rate of RuBP is reduced for large quantities of Lianas.(4) The response of chlorophyll fluorescence kinetic parameters to soil moisture stress①with the aggravation of soil moisture stress, the ration of Lianas'FV/Fm is declining, and the light translation extent is low. The decline of FV/Fm affects Photosynthetic electron transport natural progress and plant photosynthesis.②with the aggravation of soil moisture stress, the decline extent of Lianas'ΦPSⅡis increased, in turn. In the soil moisture stress of check,mild,moderate, theΦPSⅡof some Lianas could renew the primary value, but the severe water stress could limit the electron transport of reaction center in the PSⅡ,reducing the quantum yield of electron transport in the PSⅡ.③with the aggravation of soil moisture stress, the photoinhibition phenomenon is strengthen, accompanying the increscent of Lianas'NPQ, and heat dissipateion ability is also strength.④with the aggravation of soil moisture stress, the daily mean of ETR is reduced , and the descending extent is increscent, in turn. The aggravation of soil moisture stress could reduce the Photosynthetic electron transport rate. In a certain, it could not consume superfluous light energy, and induce the breach of photosynthetic organ accompanying with the aggravation of soil moisture stress and intense light.(5) The fitting soil moisture threshold value and the effective range of photosynthetic active rate for maintaining different Lianas'high-efficiency water use efficiency and natural growth①In order to maintain high-efficiency physiological using water and high-level photosynthetic productivity of Campsis radicans Seen, the fitting mass water content (Wm)is about 13.5%~19.4%, relative water content (Wr) is about 49.5%～71.1%, and the fitting photosynthetic active radiation (PAR) is in the range of 800~1600μmol·m-2·s-1. and the ordinal soil moisture content and PAR of other Lianas, as follows:②For Wisteria sinensis Sweet , the fitting Wm is about 13.1%~22.6%, Wr is about 46.5%～80.1%, and the fitting PAR is in the range of 800~1600μmol·m-2·s-1.③For Hedera neaplensis var. Sinensis , the fitting Wm is about 17.6%~20.5%, Wr is about 63.8%～74.3%, and the fitting PAR is in the range of 600~1400μmol·m-2·s-1.④For Ampelopsis brevipedunculata , the fitting Wm is about 8.9%~19.8%, Wr is about 30.8%～68.5%, and the fitting PAR is in the range of 1000~1200μmol·m-2·s-1.⑤For Parthenocissus quinquefolia , the fitting Wm is about 9.7%~14.0%, Wr is about 33.7%～48.7%, and the fitting PAR is in the range of 600~1200μmol·m-2·s-1.⑥For Parthenocissus semicordata , the fitting Wm is about 8.9%~15.7%, Wr is about 28.6%～50.2%, and the fitting PAR is in the range of 400~800μmol·m-2·s-1.⑦For Euonymus fortunei , the fitting Wm is about 9.8%~15.2%, Wr is about 34.0%～52.7%, and the fitting PAR is in the range of 600~1000μmol·m-2·s-1.⑧For E.f.var.minimus Rhed , the fitting Wm is about 8.2%~18.3%, Wr is about 32.3%～72.2%, and the fitting PAR is in the range of 600~1200μmol·m-2·s-1.⑨For Euonymus scandens Graham , the fitting Wm is about 11.2%~21.2%, Wr is about 41.0%～77.7%, and the fitting PAR is in the range of 600~1000μmol·m-2·s-1.⑩For Lonicera japonica Thunb.cv.tomentosa , the fitting Wm is about 11.4%~19.8%, Wr is about 40.4%～70.3%, and the fitting PAR is in the range of 800~1400μmol·m-2·s-1.○11 For Lonicera japonica Thunb.var , the fitting Wm is about 9.0%~17.2%, Wr is about 31.6%～60.5%, and the fitting PAR is in the range of 600~1200μmol·m-2·s-1.○12 For Lonicera japonica Thunb.cv. Variegatum , the fitting Wm is about 12.9%~17.9%, Wr is about 46.2%～64.2%, and the fitting PAR is in the range of 800~1200μmol·m-2·s-1.○13 For Lonicera japonica Thunb.cv.meng , the fitting Wm is about 12.0%~16.4%, Wr is about 43.3%～59.2%, and the fitting PAR is in the range of 600~1400μmol·m-2·s-1.○14 For Lonicera japonica Thunb.cv.ungulata , the fitting Wm is about 17.3%~22.3%, Wr is about 60.7%～78.2%, and the fitting PAR is in the range of 1000~1400μmol·m-2·s-1. (6) The characters of soil moisture adaptation and light adaptation for Lianas ①L. j. cv.ungulata,E. scandens,E. fortunei,L. j. cv.meng, et al Lianas belong to typical sun plants. A. brevipedunculata,P. quinquefolia,H. neaplensis var, et al Lianas belong to general photophilous plants. C. radicans,L. j. cv.tomentosa,W. sinensis,L. j. var , et al Lianas belong to helioskiophytes . E.f.var.minimus Rhed,P. semicordata,L. j.cv. Variegatum , et al Lianas belong to typical shade plants.②In the precondition of maintaining high Pn, the order of waterlogging tolerance is E.f.var.minimus Rhed> W. sinensis > L. j. cv.ungulata > E. scandens > H. neaplensis var> P. semicordata> L. j. var> P. quinquefolia > E. fortunei > C. radicans > L. j. cv.tomentosa > A. brevipedunculata > L. j.cv. Variegatum > L. j. cv.meng . the order of drought tolerance is > P. semicordata > A. brevipedunculata > L. j. var > E.f.var.minimus Rhed > P. quinquefolia > E. fortunei > L. j. cv.tomentosa > E. scandens > L. j. cv.meng > L. j.cv. Variegatum > W. sinensis > C. radicans > L. j. cv.ungulata > H. neaplensis var.
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