| Soil nutrients in the natural environment showed different distribution,it’s called heterogeneity.Whether clonal plants or non-clonal plants can adapt to this heterogeneity by altering root length,weight,density and the absorption rate of water,respond to the survival pressure caused by heterogeneous environments.Different from non-cloned plants,clonal plants can adapt to the survival pressure from heterogeneous environment through physiological integration,which can improve the survival probability.It is instructive for the cultivation of turfgrass and provide theoretical basis for turfgrass survival in heterogeneous soil to study the ecological adaptability of turfgrass in heterogeneous soil.The experiment used perennial Zoysia japonica as research materials,which were set into connected and disconnected groups and were added different nutrient to simulate the heterogeneous environment of soil nutrients in nature.The roots weight,the root water content,nitrogen,phosphorus,and potassium content changes in the different treatments were analyzed to study the adaptability of Zoysia japonica in heterogeneous soils.The main results are as follows:1.The dry weight and fresh weight of Zoysia japonica clonal ramets roots was the same change under the conditions of soil nutrient heterogeneity.The plasticity response pattern of the root system of Zoysia japonica showed that the weight of the ramets roots under low nutrition conditions were higher than the ramets roots under medium and high nutrition conditions under nutrient heterogeneity.Corresponding to the morphological characteristics of the root system that the ramets increase the investment of the root system under the low nutrient,and the ramets decrease the investment of the root system under the medium and high nutrient;There exists integration between the mother ramet in the medium and high nutrient and the daughter ramet in the low nutrient.Integration can reduce the investment of daughter ramet root system,The daughter ramet are more adapted to the poor environment.Because of the Zoysia japonica roots dry weight and fresh weight change werebasically same,the change of moisture content is relatively gradual.2.Under nutrient heterogeneity,Zoysia japonica in order to adapt to heterogeneity soil will occur nitrogen physiological integration,in medium and high nutrient ramet transferred a small part of ammonium nitrogen to the plants root in low nutrient,and more ammonium nitrogen were be transferred to the top;For the nitrate nitrogen of the ramet roots,mother ramet in the high nutrient transfered ammonium nitrogen to the daughter ramet in low nutrient,the daughter ramet in the high nutrient did not transfer ammonium nitrogen to the mother ramet in low nutrient;For the total nitrogen of the ramet root,the ramet in high nutrient transmitted nitrogen to the ramet in low nutrient,but the overall transmission degree was that the transmission capacity of the mother ramet to the daughter ramet was greater than the daughter ramet to the mother ramet,the transmission intensity of the total nitrogen to the top was greater than to the base.3.Zoysia japonica root system has the greatest absorption of phosphorus in medium nutrient,was the most suitable for growth.the absorption effect of phosphorus in high nutrient is not obvious.Under heterogeneity nutrient,phosphorus was transferred to the ramet in low nutrient through the stems of Zoysia japonica.That is,physiological integration had a bidirectional nature,but the transmission degree was that the transmission capacity of the mother ramet to the daughter ramet was greater than that the daughter ramet to the mother ramet.4.Under heterogeneity nutrient,The distribution of potassium content in Zoysia japonica root system was related to the soil nutrient content.The potassium content in the root system which under the low nutrient soil was lower than that under medium and high nutrient.In general,potassium was in a stable content.On the one hand,it may be that the potassium was transported faster,and was used for the growth of its own ramets in the aerial parts.On the other hand,potassium was transported to the aerial part,one part was used for the growth of its own ramets,and a part was transmitted through stems to another part. |