Study On The Evolutionary Structure Of Tilia Amurensis Rupr.in Different Ecological Environments

Tilia amurensis Rupr.is growing in shade environment and sun environment as experimental materials,by applicating the technology of paraffin slicing,apparent structural production of plants,wood slicing and wood segregation to have a comparative study on the anatomical structure of caulicle,petiole,leaf,wood,and leaf plant print characteristics of experimental material in two different environments.Microphotography and SPSS 20.0 software was used for mathematical statistics and analysis of experimental data.The experimental results show that,the same plant can resist different environments and evolve different structures,the external morphology and internal anatomy of T.amurensis Rupr.in the two environments are different.The results of a series of experiments fully prove that the evolution of plant structure is closely related to its environment.Under the shade environment,its outer tangential wall cuticle is thinner,the aerenchyma and secretory cavity are developed,but the vascular bundle is underdeveloped.The cuticle of the petiole is thinner,pore chain of vascular bundle xylem is shorter,and there is no pipe hole group,so the capability of transporting is weak.Leaf epidermal cell anticlinal wall is type F,hypodermis' stoma is protuberance but it's fewer,all of those are the representative evolution structure of the shade environment.The palisade tissue's layer is less,but the spongy tissue is well developed.There have secretory cells in ray cells and bast fiber.The wood is ring-porous,pore chain is shorter and less.The vessel element which with spiral thickening is single punch,bore diameter is smaller,and its sidewall striate hole is each column type,all of those are the evolutionary structural features in order to resist the shade environment.Under the sun environment,it's corneous layer of caulicle is thicker,collenchymas of exoderm are about 5-6 layers,and the vascular bundle is developed,there have secretory organization which are shaped of plum blossom in the pith,and there is a large number of plasmodesmata between cells,which strengthen the transport efficiency of organic matter.The caulicle which with double rays,its vascular tissue of is developed,the xylem pore cluster is more,which increases the ability of water and inorganic salts.The vascular bundle of pith almost occupy the whole position of the pith and replaced it,which enhances transport capacity.The cuticle of the petiole is thicker and vascular tissue is developed,the vascular tissue which with vascular group is developed,its pore chain is longer,so the capability of transporting is strong.The leaf cuticle is thicker,leaf epidermal which with secretory cells,its degree of specialization is high,and the leaf epidermal cell anticlinal wall is also type F.The palisade organization is developed and the spongy parenchyma is dense.There has the distribution of secretory cell and crystals in leaf.Wood is ring-porous,its pipe hole chain is longer and more.The xylem vascular with spiral thickening is single punch,and bore diameter is larger,its sidewall striate hole is each column type,the aperture is bigger.all of those are the evolutionary structural features in order to resist the sun environment.From what has been discussed above,the evolvement structure of T.amurensis Rupr.in different ecological environment is closely related to the environment in which it grows,it is the result of the long-term adaptation to its living environment.To some extent,it accurately proves the theory of ecological environment can influence the plant morphology.This paper provides the first-hand theoretical basis for selection breeding and rational exploitation of the T.amurensis Rupr.,in addition,it provides an important reference for the first-hand experimental data of research on resistance and bee research of the T.amurensis Rupr.